The film opens with a prologue scene set in a 19th-century Eastern European shtetl. In the scene, filmed entirely in Yiddish, Velvel (Allen Lewis Rickman) comes home during a snowstorm and tells his wife (Yelena Shmulenson) that he had been helped along his way by an acquaintance of hers, Reb Groshkover (Fyvush Finkel), whom he has invited in for soup. However, the wife informs him that the Reb had died three years prior, and that this visitor must be a dybbuk, an undead being of Jewish folklore. The guest laughs off this suggestion, but eventually she stabs the visitor, and he goes back out to the snow, leaving the mystery unresolved.
The remainder of the story takes place in the Twin Cities in 1967, where Jewish professor of physics and family man, Larry Gopnik (Michael Stuhlbarg) lives with his troubled family. His son Danny (Aaron Wolff) habitually smokes marijuana. Larry's wife Judith (Sari Lennick) wants a divorce so she can be with family friend and widower, Sy Ableman (Fred Melamed). Daughter Sarah (Jessica McManus) appears to be stealing money from her father to get a nose job, and his brother Arthur (Richard Kind) has been crashing on their couch for several months writing a dense numerological treatise.
At Judith and Sy's insistence, Larry and Arthur move to a nearby motel, after which Judith empties their joint financial accounts, leaving Larry penniless. One of Larry's students, Clive, claims his exams are unfair; after he leaves, Larry finds an envelope stuffed with thousands of dollars, and is furious that Clive would try to bribe him. Larry is up for tenure, but the committee has received anonymous defamatory letters about him. Soon after, Sy is killed in a car accident and it's revealed that he was penniless, saddling Larry with Sy's funeral expenses.
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